168 Works

Revision of Agraptocoris Reuter (Heteroptera: Miridae: Phylinae), with description of five new species and a review of aedeagal terminology — Electronic Supplement File 1

Fedor V. Konstantinov
Electronic Supplement File 1: konstantinov-agraptocorismiridae-asp2019-electronic-supplement-1.nex — Matrix for morphological characters.

The first fossil free-living late instar larva of Strepsiptera (Insecta) — Electronic Supplement File 3

Hans Pohl, Jörg U. Hammel, Adrian Richter & Rolf G. Beutel
Electronic Supplement File 3: pohl&al-strepsipterafossillarva-asp2019-electronicsupplement-3.nex — Data matrix with characters of males, females and immature stages (fossil larva coded as female).

Revised phylogenetic relationships within the Drosophila buzzatii species cluster (Diptera: Drosophilidae: Drosophila repleta group) using genomic data

Juan Hurtado, Francisca Almeida, Santiago Revale & Esteban Hasson
The Drosophila buzzatii cluster is a South American clade that encompasses seven closely related cactophilic species and constitutes a valuable model system for evolutionary research. Though the monophyly of the cluster is strongly supported by molecular, cytological and morphological evidence, phylogenetic relationships within it are still controversial. The phylogeny of the D. buzzatii cluster has been addressed using limited sets of molecular markers, namely a few nuclear and mitochondrial genes, and the sharing of fixed chromosomal...

The genetic identity of the only Italian population of the genus Macroprotodon Guichenot, 1850 on the island of Lampedusa, Sicily

Francesco Paolo Faraone, Raffaella Melfi, Matteo Riccardo Di Nicola, Gabriele Giacalone & Mario Lo Valvo
The only Italian population of false smooth snakes is found on Lampedusa, a small island located in the Sicilian Channel and part of the African continental shelf. The taxonomic identity of this population is currently uncertain, although it is most often attributed to Macroprotodon cucullatus textilis on a morphological basis. We present here the first genetic data on this population. The analysis carried out on the mitochondrial cytochrome b gene shows that the Lampedusan false...

Knockdown of zax in Xenopus laevis leads to craniofacial malformations and the absence of the intramandibular joint

Paul Lukas, Jennifer Schmidt & Lennart Olsson
The jaws of anuran tadpoles consist of several evolutionary novelties, which are unique within the vertebrates. The most noticeable of these novelties are two cartilaginous elements called rostralia and the derived organisation of cranial muscles, which form an entirely new feeding apparatus. The suprarostral cartilage is located in the upper and the infrarostral cartilages in the lower jaw. Between infrarostral and Meckel’s cartilage an intramandibular joint is present. The morphological diversity of anuran tadpoles and...

The phylogeny of Ptiliidae (Coleoptera: Staphylinoidea) – the smallest beetles and their evolutionary transformations

Alexey A. Polilov, Ignacio Ribera, Margarita I. Yavorskaya, Anabela Cardoso, Vasily V. Grebennikov & Rolf G. Beutel
The smallest beetles and the smallest non-parasitic insects belong to the staphylinoid family Ptiliidae. Their adult body length can be as small as 0.325 mm and is generally smaller than 1 mm. Here we address the phylogenetic relationships within the family using formal analyses of adult morphological characters and molecular data, and also a combination of both for the first time. Strongly supported clades are Ptiliidae + Hydraenidae, Ptiliidae, Ptiliidae excl. Nossidium, Motschulskium and Sindosium,...

Taxonomic revision and comments on two groups of the genus Coniophanes (Squamata: Dipsadidae)

Ricardo Palacios-Aguilar & Oscar Flores-Villela
A revision of the Coniophanes lateritius and C. piceivittis groups was conducted to evaluate the taxonomic status of their members. The supraspecific groups of Coniophanes can be easily distinguished from each other, but the species within them exhibit wide overlap in scutellation. Apparently, these taxa can be differentiated by color pattern and geographic distribution. However, we report polymorphism in the color pattern of the lateritius group. Maxillary and hemipenial morphology can be useful and informative...

Psilorhynchus kamengensis, a new species of fish (Teleostei: Psilorhynchidae) from northeast India

Abhinit Dey, Hrishikesh Choudhury, Abhishek Mazumder, Ratul Ch. Bharali, Sarbojit Thaosen & Dandadhar Sarma
Psilorhynchus kamengensis, new species, is described from a stream of the Kameng River, a northern tributary to the Brahmaputra drainage, Arunachal Pradesh, northeast India. The new species belongs to the P. balitora species group and can be distinguished from all other members of this group by the presence of a longer post-epiphyseal fontanelle, 3 unbranched anal-fin rays, 9 + 8 caudal-fin rays, 35 vertebrae and fin coloration. The validity of the species is also corroborated by molecular...

Turning to the dark side: Evolutionary history and mole­cular species delimitation of a troglomorphic lineage of armoured harvestman (Opiliones: Stygnopsidae)

Jesús A. Cruz-López, Rodrigo Monjaraz-Ruedas & Oscar F. Francke

Turning to the dark side: Evolutionary history and mole­cular species delimitation of a troglomorphic lineage of armoured harvestman (Opiliones: Stygnopsidae)

Jesús A. Cruz-López, Rodrigo Monjaraz-Ruedas & Oscar F. Francke

Revisited phylogeny of Scleroderminae (Hymenoptera: Bethylidae) reveals a plastic evolutionary history

Juan M. Vargas R., Wesley D. Colombo & Celso O. Azevedo

The African-Iberian connection in Odonata: mtDNA and ncDNA based phylogeography of Aeshna cyanea (Müller, 1764) (Odonata: Aeshnidae) in Western Palaearctic

Thomas J. Simonsen, Kent Olsen & Marie Djernæs
We explore the phylogeography and inter-population relationships of the Southern Hawker dragonfly, Aeshna cyanea (Müller) in the Western Palaearctic region based on 603 bp Cytochrome Oxidase Subunit 1 (COI) mtDNA and 732 bp Internal Transcribed Spacer region (Internal Transcribed Spacer 1, 5.8S ribosomal RNA gene and Internal Transcribed Spacer 2, ITS region) ncDNA with an increased sampling from Europe compared to a previous study. Both DNA fragments recover a remarkable and compatible pattern: the recently...

The African-Iberian connection in Odonata: mtDNA and ncDNA based phylogeography of Aeshna cyanea (Müller, 1764) (Odonata: Aeshnidae) in Western Palaearctic – Electronic Supplements

Thomas J. Simonsen, Kent Olsen & Marie Djernæs

Cryptic diversity in the whip spider genus Paraphrynus (Amblypygi: Phrynidae): integrating morphology, karyotype and DNA – Electronic Supplements

Michael Seiter, Azucena C. Reyes Lerma, Jiří Král, Alexandr Sember, Klára Divišová, José G. Palacios-Vargas, Pío A. Colmenares, Stephanie F. Loria & Lorenzo Prendini

Phylogeny and Systematics of Wood-Degrading Neotro­pical Bess Beetles (Coleoptera: Passalidae: Passalinae)

Cristian Fernando Beza-Beza, Larry Jiménez-Ferbans & Duane D. McKenna
Passalidae (Coleoptera: Scarabaeoidea) are saproxylophagous beetles that live and feed inside decaying wood. Passalid diversity and biogeography are relatively well-established at the tribal-level, particularly in the New World. However, a robust phylogenetic framework to test the current taxonomic status of the generic groups in the family is lacking. In the New World, Passalidae is represented by the tribes Passalini and Proculini. To test the monophyly of Proculini and Passalini and reconstruct relationships among the New...

DNA barcoding and molecular taxonomy of dark-footed forest shrew Myosorex cafer in the Eastern Cape and KwaZulu-Natal, South Africa

Emmanuel Matamba, Leigh R. Richards, Michael I. Cherry & Ramugondo V. Rambau
There is a paucity of molecular DNA barcoding informatics on the South African fauna, particularly on terrestrial small mammals. This study tested the utility of DNA barcoding in the dark-footed forest shrew (Myosorex cafer) from forested regions of the Eastern Cape and KwaZulu-Natal provinces of South Africa. Sampled forests included coastal scarp, dune forests and inland Afromontane mistbelt forests. Sequences of mtDNA cytochrome oxidase subunit I (COI, 623 bp), were generated for a total of...

Phylogeny of the Neotropical genus Stenosigma Giordani Soika, 1978 (Hymenoptera, Vespidae, Eumeninae) based on morphological data

Wellington Donizet Ferreira & Marcel Gustavo Hermes
Stenosigma Giordani Soika, 1990 is a small Neotropical genus of potter wasps (Hymenoptera, Vespidae, Eumeninae), with an interesting distributional pattern and a recent revision published. Unfortunately, the genus lacks a phylogenetic hypothesis with all species included — one major impediment for future biogeographic studies and hypotheses for evolutionary scenarios. Therefore, a phylogenetic treatment would help enlighten the history of this Neotropical wasp genus. We present the first phylogeny of Stenosigma based on cladistic methods using...

Kruia rediscovered: phylogenetic implications, larval morphology, and biology of an enigmatic hydrophilid beetle from western Africa (Coleoptera: Hydrophilidae)

Martin Fikáček, Qing-Hao Zhao, Ishmeal N. Kobe & Vasily Grebennikov
The enigmatic terrestrial water scavenger beetle (Hydrophilidae: Sphaeridiinae: Coelostomatini) of the genus Kruia Spangler & Perkins, 1981 was known by a single specimen from Liberia. We rediscovered it at two forested localities in Cameroon (Mt. Cameroon and Mt. Kupe, both of volcanic origin). Adults and larvae of a new species were collected by sifting forest leaf litter. To reveal the phylogenetic position of Kruia, we analyzed a dataset comprising 32 species (incl. 29 of the...

New insight into the systematics of Heteromurini (Collembola: Entomobryidae: Heteromurinae) with special reference to Alloscopus and Sinodicranocentrus gen.n.

Feng Zhang, Nikolas G. Cipola, Zhi-Xiang Pan & Yinhuan Ding
Classification of Heteromurini represented by Heteromurus and Dicranocentrus is currently under debate due to the presence of 5 or 6 antennal segments, respectively. Among the Heteromurini, Alloscopus is traditionally considered to be closely related to Heteromurus but its phylogenetic position is challenged by recent molecular evidence. A multilocus molecular phylogeny and topology tests conducted here indicate the closer relationship of Alloscopus and Dicranocentrus, and one non-Dicranocentrus basal group, Sinodicranocentrus gen.n. Two new species, Alloscopus bannaensis...

Phylogeny of the genus Tephritis Latreille, 1804 (Diptera: Tephritidae)

Severyn V. Korneyev, John T. Smit, Daniel L. Hulbert, Allen L. Norrbom, Stephen D. Gaimari, Valery A. Korneyev & James J. Smith
The phylogenetic relationships of Tephritis Latreille, one of the largest genera of true fruit flies, are poorly understood due to the large number of species, lack of reliable morphological characters and difficulties in identification. In the present study we used two datasets to address these problems: one with 35 species of the genus Tephritis (28 from the Palaearctic and seven from the Nearctic Region) and seven species from closely related genera of the tribe Tephritini...

Phylogeny and life history evolution of Blaberoidea (Blattodea)

Marie Djernæs, Zuzana Kotyková Varadínová, Michael Kotyk, Ute Eulitz & Klaus-Dieter Klass
Blaberoidea, comprised of Ectobiidae and Blaberidae, is the most speciose cockroach clade and exhibits immense variation in life history strategies. We analysed the phylogeny of Blaberoidea using four mitochondrial and three nuclear genes from 99 blaberoid taxa. Blaberoidea (excl. Anaplectidae) and Blaberidae were recovered as monophyletic, but Ectobiidae was not; Attaphilinae is deeply subordinate in Blattellinae and herein abandoned. Our results, together with those from other recent phylogenetic studies, show that the structuring of Blaberoidea...

Revision of Carexomyza Roháček with descriptions of three new Nearctic species (Diptera: Anthomyzidae)

Kevin N. Barber & Jindřich Roháček
The formerly monotypic genus Carexomyza Roháček, 2009 (Diptera: Anthomyzidae) is revised to include one Palaearctic, viz. C. caricis (Roháček, 1999), and three new Nearctic species, viz. C. lonsdalei sp.n., C. marshalli sp.n. and C. browni sp.n., all from Canada and the northern United States of America. Carexomyza is re-diagnosed and its phylogenetic position within extant Anthomyzinae discussed. All species are described or diagnosed and illustrated and a hypothesis about their relationships (based both on morphological...

Kruia rediscovered: phylogenetic implications, larval morphology, and biology of an enigmatic hydrophilid beetle from western Africa (Coleoptera: Hydrophilidae) – Electronic Supplement

Martin Fikáček, Qing-Hao Zhao, Ishmeal N. Kobe & Vasily Grebennikov
Supplementary file: fikacek&al-kruia-electronic-supplement.doc. — Table S1. Amplification programs and primers. — Table S2. PartionFinder2 output: IQ-Tree block and MrBayes block for dataset used. — Fig. S1. Maximum likelihood phylogenetic tree of the Coelostomatini. — Fig. S2. Bayesian analysis phylogenetic tree of the Coelostomatini.

Short note to the inoceramid bivalve Inoceramus pictus concentricoundulatus Tröger, 1967 – holotype and further specimens from the lower Upper Cenomanian of Saxony (Germany)

Birgit Niebuhr
To date, there is a lack of clarity in the literature due to the holotype of the bivalve subspecies Inoceramus pictus concentricoundulatus Tröger, 1967. The specimen of the first erection of the subspecies name by Tröger (1967: 46, pl. 3, fig. 7) from the lower Upper Cenomanian of Saxony, Germany, still exists in the TU Bergakademie Freiberg (holotye FG: 253/379). This I. pictus subspecies rarely occurs; six more specimens are hosted in the Museum für...

Wolfram aus dem sächsischen Erzgebirge

Heinz Schulz †
Nach von England ausgehender Verwendung von Wolfram begann um 1860 im Osterzgebirge die Nutzung vordem in Versatz oder Schlacken verbliebener Wolframinhalte. Im Raum Zinnwald erlangte Wolframit rasch wirtschaftliche Bedeutung und sicherte den Erhalt der Bergbaubetriebe. Mit der Entwicklung von Wolframerzeugnissen für die Metallbearbeitung sowie von Glühfäden in Leuchtmitteln stieg der Bedarf und erreichte im Ersten Weltkrieges einen ersten Höhepunkt. Die Folge war der Aufschluss weiterer Vorkommen im Westerzgebirge und die Gewinnung von Wolfram aus Schlacken...

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