Extending the footprint record of Pareiasauromorpha to the Cisuralian: earlier appearance and wider palaeobiogeography of the group

Lorenzo Marchetti, Sebastian Voigt, Eudald Mujal, Spencer Lucas, Heitor Francischini, Josep Fortuny & Vincent Santucci
Pareiasauromorpha is one of the most important tetrapod groups of the Permian. Skeletal evidence suggests a late Kungurian origin in North America, whereas the majority of occurrences come from the Guadalupian and Lopingian of South Africa and Russia. Pareiasauromorpha footprints include the ichnogenus Pachypes, that is, however, unknown from strata older than late Guadalupian. A revision of several Pachypes-like footprints from the Cisuralian–Guadalupian of Europe and North America confirm the occurrence of this ichnogenus and...
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