Social insect colonies are more likely to accept unrelated queens when they come with workers

Ornela De Gasperin, Pierre Blacher & Michel Chapuisat
Relatedness underlies the evolution of reproductive altruism, yet eusocial insect colonies occasionally accept unrelated reproductive queens. Why would workers living in colonies with related queens accept unrelated ones, when they do not gain indirect fitness through their reproduction? To understand this seemingly paradox, we investigated whether acceptance of unrelated queens by workers is an incidental phenomenon resulting from failure to recognize non-nestmate queens, or whether it is adaptively favored in contexts where cooperation is preferable...
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3 downloads reported since publication in 2021.

These counts follow the COUNTER Code of Practice, meaning that Internet robots and repeats within a certain time frame are excluded.
What does this mean?