Genetic divergence, admixture and subspecific boundaries in a peripheral population of great tit, Parus major (Aves, Paridae)

Sahar Javaheri Tehrani, Laura Kvist, Omid Mirshamsi, Seyed Mahmoud Ghasempouri & Mansour Aliabadian
Secondary contact zones have been formed between several pairs of avian species and subspecies in northern and northeastern Iran during the post-Pleistocene and Holocene times. Three subspecies groups out of the four in the great tit (Parus major), namely, major, bokharensis and cinereus are supposed to have come into local or regional secondary contact in northeastern Iran. Parus major intermedius known from this region, has long been suggested to have a hybrid origin, stemming from...
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These counts follow the COUNTER Code of Practice, meaning that Internet robots and repeats within a certain time frame are excluded.
What does this mean?