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FIGURE 1 in Rhopalione kali sp. nov., first known epicaridean parasite on the Malaysian pinnotherid crab, Serenotheres besutensis (Serène, 1967)

Shane T. Ahyong & Christopher B. Boyko
FIGURE 1. Rhopalione kali sp. nov., Perhentian Besar, Malaysia: A–H, holotype female, length 4.00 mm, MNHN-IU-2013- 19983; I–L, allotype male, length 2.70 mm, MNHN-IU-2013-19984. A, I, dorsal view; B, J, ventral view; C, K, left antennule and antenna, ventral view; D, left maxilliped with lateral lobe of barbula, external view; E, barbula, median region; F, L, right pereopod 1 of female and male, respectively; G–H, left oostegite 1, external and internal views. Scale: A–B =...

FIGURE 1 in Rhopalione kali sp. nov., first known epicaridean parasite on the Malaysian pinnotherid crab, Serenotheres besutensis (Serène, 1967)

Shane T. Ahyong & Christopher B. Boyko
FIGURE 1. Rhopalione kali sp. nov., Perhentian Besar, Malaysia: A–H, holotype female, length 4.00 mm, MNHN-IU-2013- 19983; I–L, allotype male, length 2.70 mm, MNHN-IU-2013-19984. A, I, dorsal view; B, J, ventral view; C, K, left antennule and antenna, ventral view; D, left maxilliped with lateral lobe of barbula, external view; E, barbula, median region; F, L, right pereopod 1 of female and male, respectively; G–H, left oostegite 1, external and internal views. Scale: A–B =...

FIGURE 12 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 12. Map showing the currently known distribution of Achalinus juliani sp. nov. in Cao Bang Province (1 = Ha Lang, type locality, 2 = Nguyen Binh), of Achalinus timi sp. nov. in Son La Province (3 = Copia Nature Reserve), and of Achalinus emilyae sp. nov. in Quang Ninh and Bac Giang provinces (4 = Dong Son-Ky Thuong Nature Reserve, type locality, 5 = Suoi Tuyen).

FIGURE 12 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 12. Map showing the currently known distribution of Achalinus juliani sp. nov. in Cao Bang Province (1 = Ha Lang, type locality, 2 = Nguyen Binh), of Achalinus timi sp. nov. in Son La Province (3 = Copia Nature Reserve), and of Achalinus emilyae sp. nov. in Quang Ninh and Bac Giang provinces (4 = Dong Son-Ky Thuong Nature Reserve, type locality, 5 = Suoi Tuyen).

FIGURE 11 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 11. Habitat of Achalinus emilyae sp. nov. in Bac Giang Province, northern Vietnam. Phot. T. Ziegler

FIGURE 11 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 11. Habitat of Achalinus emilyae sp. nov. in Bac Giang Province, northern Vietnam. Phot. T. Ziegler

FIGURE 10 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 10. Portrait of the paratype of Achalinus emilyae sp. nov. (VNMN 1334) from Suoi Tuyen, Bac Giang Province, northern Vietnam. Phot. T. Ziegler

FIGURE 10 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 10. Portrait of the paratype of Achalinus emilyae sp. nov. (VNMN 1334) from Suoi Tuyen, Bac Giang Province, northern Vietnam. Phot. T. Ziegler

FIGURE 9 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 9. Paratype of Achalinus emilyae sp. nov. (VNMN 1334) from Suoi Tuyen, Bac Giang Province, northern Vietnam, in life. Phot. T. Ziegler

FIGURE 9 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 9. Paratype of Achalinus emilyae sp. nov. (VNMN 1334) from Suoi Tuyen, Bac Giang Province, northern Vietnam, in life. Phot. T. Ziegler

FIGURE 8 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 8. Holotype of Achalinus emilyae sp. nov. (IEBR 4465) from Dong Son-Ky Thuong Nature Reserve, Hoanh Bo District, Quang Ninh Province, northern Vietnam, in life. Phot. M. van Schingen

FIGURE 8 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 8. Holotype of Achalinus emilyae sp. nov. (IEBR 4465) from Dong Son-Ky Thuong Nature Reserve, Hoanh Bo District, Quang Ninh Province, northern Vietnam, in life. Phot. M. van Schingen

FIGURE 7 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 7. Midbody region of the preserved holotype of Achalinus timi sp. nov. (IEBR A.2018.10) in dorsal (top) and ventral view (bottom). Phot. T. Ziegler

FIGURE 7 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 7. Midbody region of the preserved holotype of Achalinus timi sp. nov. (IEBR A.2018.10) in dorsal (top) and ventral view (bottom). Phot. T. Ziegler

FIGURE 6 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 6. Portrait of the preserved holotype of Achalinus timi sp. nov. (IEBR A.2018.10) from Thuan Chau District, Son La Province, northern Vietnam. Phot. T. Ziegler

FIGURE 6 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 6. Portrait of the preserved holotype of Achalinus timi sp. nov. (IEBR A.2018.10) from Thuan Chau District, Son La Province, northern Vietnam. Phot. T. Ziegler

FIGURE 5 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 5. Preserved paratypes of Achalinus juliani sp. nov. (IEBR A.2018.9, left, and VNMN 06924, right), both from Nguyen Binh District, Cao Bang Province, northern Vietnam. Phot. T. Ziegler and T. T. Nguyen

FIGURE 5 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 5. Preserved paratypes of Achalinus juliani sp. nov. (IEBR A.2018.9, left, and VNMN 06924, right), both from Nguyen Binh District, Cao Bang Province, northern Vietnam. Phot. T. Ziegler and T. T. Nguyen

FIGURE 4 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 4. Lateral view of the midbody region of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8). Phot. T. Ziegler

FIGURE 4 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 4. Lateral view of the midbody region of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8). Phot. T. Ziegler

FIGURE 3 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 3. Overall view of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8). Phot. T. Ziegler

FIGURE 3 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 3. Overall view of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8). Phot. T. Ziegler

FIGURE 2 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 2. Head views of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8) from Ha Lang District, Cao Bang Province, northern Vietnam. Phot. T. Ziegler

FIGURE 2 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 2. Head views of the preserved holotype of Achalinus juliani sp. nov. (IEBR A.2018.8) from Ha Lang District, Cao Bang Province, northern Vietnam. Phot. T. Ziegler

FIGURE 1 in Three new species of the snake genus Achalinus from Vietnam (Squamata: Xenodermatidae)

Thomas Ziegler, Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Van Pham, Mona Van Schingen, Tham Thi Nguyen & Minh Duc Le
FIGURE 1. Phylogram based on the Bayesian analysis. Numbers above and below branches are MP/ML bootstrap values and combined Bayesian posterior probabilities (>50%), respectively. Asterisk represents 100% value, and dash indicates <50% value. Scale bar shows substitutions/site.

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